Prevalence of naturally occurring natural enemies

Natural enemies (parasitoids and predators) have frequently been found to be associated with pests in a wide variety field studies on stored product insects and mites as shown by the following 171 studies (one in 1897, 4-21/decade between 1911 and 1980, and 18-34/decade between 1981 and 2020) and in a few cases, numbers of natural enemies were sufficient to reduce pest populations. For 38 natural enemy species, natural occurrence has been reported more than 4 times (Table Frequencies of natural occurrences) and 137 species were only reported once with a total of 218 species reported as naturally occurring. Studies differed in the combinations of geographic location, commodity, host species and natural enemy species. In addition to 840 reported occurrences, 1064 associations of natural enemies with commodities have been reported for 169 species (Table Commodity-Natural Enemy Associations).

Hagstrum, D. W., T. Z. Klejdysz, Bh. Subramanyam and J. Nawrot. 2013. Atlas of Stored-Product Insects and Mites. AACC International, St. Paul, Minnesota.

Hagstrum, D. W., and Bh. Subramanyam. 2009. Stored-Product Insect Resource. AACC International, St. Paul, Minnesota.

Other pages on biological control are:

Biological Control

Blattisocius tarsalis prey

Cheyletus prey

Conservation biological control for stored product insects

Insecticide Resistant Natural Enemies

Natural enemies of Tribolium castaneum

Phthorimaea operculella natural enemies

Trichogramma natural enemies of stored-product insects

References:

Abdel-Rahman, H. A., N. F. Shaumar, Z. A. Soliman, and M. A. El-Agoze 1981. Survey and Taxonomy of Parasites and Predators of Stored Grain and Grain Products Insects. Bulletin de La Societe Entomologique d’Egypte 61: 53–74. Amphibolus venator, Anisopteromalus calandrae, Bracon hebetor, Chelonus sulcatus, Coranus aegyptius, Diadegma armillata, Diadegma chrysostictos, Dibrachys cavus, Dinarmus laticeps, Habrocytus cerealellae, Holepyris hawaiiensis, Joppeicus paradoxus, Holepyris sylvanidis (Rhabdepyris zeae), Saprinus metallescens, Saprinus semistriatus, Scenopinus fenestralis, Scenopinus glabrifrons, Tenebroides mauritanicus, Theocolax elegans (Chaetospila elegans), Trichogramma evanescens, Venturia canescens

Al-Nasser, Abir Sulaiman 2011. Quantitative survey of stored products mites infesting wheat flour in Jeddah Governorate. Journal of Entomology and Nematology 3(6): 78-84. Acarophenax tribolii, Blattisocius tarsalis, Blattisocius keegani, Cheyletus malaccensis

Antoine, S., D. Clementine, O.A. Patoin and H. Jacques 2005. Field occurrence of bruchid pests of cowpea and associated parasitoids in a subhumid zone of Burkina Faso: importance on the infestation of two cowpea varieties at harvest. Plant Pathol. J. 4(1): 14-20. Dinarmus basalis

Aitken, A. D. 1975. Insect Travelers. Volume I. Coleoptera. A Survey of the Beetles Recorded from Imported Cargoes by the Insect Inspectorate of the Ministry of Agriculture, Fisheries and Food with a Special Analysis for the Years 1957 to 1969 Technical Bull. 31. Aglenus brunneus, Aprostoma planifrons, Bitoma siccana, Carcinops pumilio, Carcinops troglodytes, Colobicus parilis, Colobicus specialis, Cylidrus fasciatus, Exkorynetes analis (Corynetes analis), Hister major, Microprius confuses, Necrobia ruficollis, Necrobia rufipes, Necrobia violacaea, Opilo domesticus, Saprinus chalcites, Tarsosternus univittatus, Teretrius punctulatus, Thaneroclerus buqueti, Tillus notatus, Tyrtaeus dobsoni

Aitken, A. D. 1984. Insect Travelers. Volume II. A Survey of the Beetles Recorded from Imported Cargoes by the Insect Inspectorate of the Ministry of Agriculture, Fisheries and Food during the Period 1957 to 1977. Ministery of Agriculture, Fisheries and Food Reference Book 437. Amphibolus venator, Anisopteromalus calandrae, Antrocephalus mitys (Antrocephalus mahensis), Apanteles carpatus, Apanteles trachalus, Cephalonomia tarsalis, C. waterstoni, Cerocephala dinoderi, Diadegma chrysostictos (Angitia chrysosticta, Horogenes chrysostictos), Dinarmus basalis (Dinarmus laticeps), Dinarmus colemani, Euchalcidia caryobori, Fulvius anthocoroides (Fulcius brevicornus), Habrobracon hebetor, Holepyris glabratus (Holepyris hawaiiensis), Holepyris sylvanidis (Rhabdepyris zeae), Lariophagus distinguendus, Lyctocoris campestris, Peregrinator biannulipes (Allaeocranum biannulipes), Plastanoxus monroi, Pteromalus semotus (Habrocytus cerealella), Reduvius personatus, Theocolax elegans (Chaetospila elegans), Triaspis thoracica, Venturia canescens, Xenolytus bitinotus (Phygadens bitinotus), Xylocoris afer, Xylocoris flavipes, Xylocoris galactinus

Al-Obaidy, Shaymaa Hameed, Samar Muthanna Shahrabani, and Awatif Abdul-Fatah Hamodi 2019. New record of Anisopteromalus calandrae (Howard, 1881) (Hymenoptera: Chalcidoidea: Pteromalidae) as parasitoid of the cigarette beetle, Lasioderma serricorne (F.)(Coleoptera: Anobiidae) in Iraq. Plant Archives 19(1): 292-294.

Anonymous 1958. Control of pests in stored agricultural products with special reference to grain. Report of a survey in North and South America and certain Mediterranean countries in 1954 and 1955. Organization for European Economic Co-Operation (OEEC), Paris Anisopteromalus calandrae, Habrobracon hebetor, Venturia canescens

Arbogast, R.T., and S.R. Chini 2005. Abundance of Plodia interpunctella (Hubner) and Cadra cautella (Walker) Infesting Maize Stored on South Carolina Farms: Seasonal and Non-Seasonal Variation. Journal of Stored Products Research 41(5): 528–543. Habrobracon hebetor, Xylocoris flavipes

Arbogast, R. T., and Mullen, M. A. 1990. Interaction of maize weevil (Coleoptera: Curculionidae) and parasitoid Anisopteromalus calandrae (Hymenoptera: Pteromalidae) in a small bulk of stored corn. J. Econ. Entomol. 83(6): 2462-2468.

Arthur, F.H., D.W. Hagstrum, P.W. Flinn, C.R. Reed, and T.W. Phillips 2006. Insect Populations in Grain Residues Associated with Commercial Kansas Grain Elevators. Journal of Stored Products Research 42(2): 226–239. Anisopteromalus calandrae, Cephalonomia waterstoni, Habrobracon hebetor, Theocolax elegans (Chaetospila elegans)

Asl, Mohammad Hossein Akbari, Ali Asghar Talebi, Hashem Kamali, and Somayeh Kazemi. 2009. Stored product pests and their parasitoid wasps in Mashhad, Iran. Advances in Environmental Biology 3(3): 239-243. Anisopteromalus calandrae, Cephalonomia tarsalis, Habrobracon hebetor, Theocolax elegans, Venturia canescens

Attia, R., and A. H. Kamel. 1966. The Fauna of Stored Products in U.A.R. Bull.de La Societe Entomolique D’Egypte 49: 221–232. Amphibolus venator, Anisopteromalus calandrae (Aplastomorpha calandrae),Bracon brevicornis, Cephalonomia aculeata, Cheyletus buleri, Cheyletus eruditus, Plastanoxus munroi (Cheyletus munroi), Cheyletus trouessarti, Coranus aegyptius, Dibrachys cavus, Habrobracon hebetor, Horogenes polygona, Joppeicus paradoxus, Lariophagus distinguendus, Orius madeirensis, Meteorus rubens, Necrobia ruficollis, Necrobia rufipes, Scenopinus glabrifrons, Thaneroclerus buqueti, Theocolax elegans, Trichogramma evanescens, Trichogramma minutum, Venturia canescens, Xylocoris flavipes

Awadallah, K. T., A. I. Afifi, and I. I. El-Sebaey 1990. Population Studies on Meat Meal Insect Pests and Their Associated Natural Enemies. Bull. Soc. Entomol. Egypte 69: 159–167. Lariophagus distinguendus, Peregrinator biannulipes (Allaeocranum biannulipes)

Awadallah, K. T., M. F. S. Tawfik, and M. M. H. Abdella 1982. Population Studies on Four Insect Pests and their Natural Enemies at Four Localities of Egypt. Bull. Soc. Entomol. Egypte 64: 95–109. Habrobracon hebetor, Holepyris sylvanidis, Peregrinator biannulipes (Allaeocranum biannulipes), Venturia canescens (Nemeritis canescens), Xylocoris flavipes

Awadallah, K. T., M. F. S. Tawfik, and M. M. H. Abdella 1982. Seasonal Fluctuations of Insect Pests and their Associated Natural Enemies in Stored Barley and Sorghum at Fayoum Governorate. Egypt. Bull. Soc. Entomol. Egypt 14: 111–119. Anisopteromalus calandrae, Lariophagus distinguendus, Xylocoris flavipes

Ayyar, P. N. 1934. A very destructive pest of stored products in south India, Corcyra cephalonica, Staint. (Lep.). Bulletin of Entomological Research 25: 155–69. Antrocephalus mitys (Antrocephalus mahensis), Habrobracon hebetor

Back, E. A. 1939. The cigarette beetle as a pest of cottonseed meal. J. Econ. Entomol. 32: 739–749. Anisopteromalus calandrae

Bare, C.O. 1942. Some natural enemies of stored-tobacco insects, with biological notes. J.Econ. Entomol. 35: 185-189. Anisoptermalus calandrae, Habrobracon hebetor, Lariophagus distinguendus, Mesostenus gracilis, Theocolax elegans, Venturia canescens

Battu, G. S., S. S. Bains, and A. S. Atwal. 1975. Natural enemies of Trogoderma granarium Everts infesting wheat in the rural stores in the Punjab (India). Bulletin of Grain Technology(India) 13: 52-55. Four spiders (Pholcidae: Artema atlanta, Oecobiidae: Oecobus putus, Selenopidae: Selenops agumbensis, Uloboridae: Uloborus danolius), Theocolax elegans (Chaetospila elegans)

Belda, C., and J. Riudavets 2010. Distribution of Insect Pests and their Natural Enemies in a Barley Pile. p. 741–745. In Proceedings of the 10th International Working Conference on Stored Product Protection. Anisopteromalus calandrae, Cephalonomia waterstoni, Withius piger

Belda, Consuelo, and Jordi Riudavets 2013. Natural Enemies Associated with Lepidopteran Pests in Food and Feed Processing Companies. Journal of Stored Products Research 53: 54–60. Anisopteromalus calandrae, Cephalonomia waterstoni, Habrobracon hebetor, Venturia canescens

Belton, C. H. 1951. Note on Some Insect Pests in Stored Products. New Zealand Journal of Science and Technology 32 (1): 44. Lyctocoris campestris

Beratlief, C. 1967. An Important Parasite of Cereal Weevils Anisopteromalus calandrae How. Analele Institutului de Cercetari Pentru Protectia Plantelor 4: 253–254. 7.2-18% Sitophilus oryzae parasitized by Anisopteromalus calandrae

Bilal, H., A. M. Basheer, and A. T. Saleh. 2011. Survey and abundance of common parasitoid and predatory species of the cigarette Beetle, Lasioderma serricorne (F.) (Coleoptera: Anobiidae) in tobacco storage warehouses in Syria. Egyptian Journal of Biological Pest Control 21: 151–153. Anisopteromalus calanrae, Ericydnus sipylus, Lariophagus distinguendus, Pteromalus cereallae

Binoy, C. F. 2015. Hymenopteran Parasitoids Associated with Some Important Stored Product Pests in Kerala, India. International Journal of Science, Environment and Technology 4(2): 509–513. Anisopteromalus calandrae was found to parasitise 11 stored product pests followed by Theocolax elegans (7 pests), Lariophagus distinguendus (6 pests), Dinarmus basalis (4 pests) and Bracon hebetor (3 pests).

Bissell Theodore L. 1940 Curculionidae, Bruchidae, Lepidoptera, and their parasites, infesting the seed pods of cowpea and various wild plants. J. econ. Entomol.33: 844-847. Anisopteromalus calandrae (Aplastomorpha calandrae), Dinarmus basalis (Bruchobius laticeps), Lariophagus texanus

Boateng, B.A. 1997. Effects of maize variety and season on population dynamics of the larger grain borer, P. truncatus (Horn) and the maize weevil, Sitophilus zeamais Motschulsky, and grain losses in the traditional ‘Ewe’ storage bam in Ghana. M.Phil. Thesis. University. Anisopteromalus calandrae, Theocolax elegans

Bodlah, Imran, Muhammad Adnan Bodlah, Muhammad Naeem, Sobia Khaliq, and Ayesha Aihetasham. 2016. First record of Anisopteromalus calandrae (howard, 1881)(Hymenoptera: Chalcidoidea: Pteromalidae) as a potential bio-control agent of stored grain beans from rawalpindi and Islamabad. Asian J Agric Biol 4 (): 73-76. Detected on mung bean (Vigna radiata) and kidney bean (Phaseolus vulgaris); damaged by two species of Genus Callosobruchus: C. chinensis and C. maculatus.

Booker, R. H. 1965. List of insect species found in association with cowpeas at Samaru. Samaru Miscellaneous Paper 9 Anisopteromalus calandrae, Dinarmus laticeps parasitizing Callosobruchus maculatus

Borgemeister, C., WG Meikle, D. Scholz, C. Adda, P. Degbey, and RH Markham. 1997. Seasonal and Weather Factors Influencing the Annual Flight Cycle of Prostephanus truncatus (Coleoptera: Bostrichidae) and Its Predator Teretriosoma nigrescens (Coleoptera: Histeridae) in Benin. Bulletin of Entomological Research 87(3): 239–246.

Brower, JH. 1984. The Natural Occurrence of the Egg Parasite, Trichogramma, on Almond Moth Eggs in Peanut Storages in Georgia. Journal of the Georgia Entomological Society 19(3): 283–290. Natural populations of Trichogramma pretiosum and Trichogramma parkeri parasitized Cadra cautella eggs at 3 out of 7 peanut warehouses.

Champ, B. R. 1966. Insects and mites associated with stored products in Queensland. 3. Hymenoptera. Queensland Journal of Agricultural and Animal Sciences 23: 177–195. Anisopteromalus calandrae, Apanteles carpatus, Apanteles galleriae, Bracon gelechiae, Cephalonomia tarsalis, Cephalonomia waterstoni, Cerocephala dinoderi, Chremylus elaphus, Habrobracon hebetor, Holepyris sylvanidis, Lariophagus distinguendus, Microchelonus phthorimaea, Paralitomastix koehleri (Copidosoma koehleri), Pteromalus cerealella (Pteromalus semotus, Habrocytus cerealella), Theocolax elegans (Choetospila elegans), Venturia canescens (Devorgilla canescens)

Champ, B. R. 1966. Insects and mites associated with stored products in Queensland. 4. Acarina and Pseudoscorpiones. Queensland Journal of Agricultural and Animal Sciences 23: 197–210. Acarophenax tribolii, Blattisocius tarsalis (Melichares tarsalis), Cheletomorpha lepidoptorum, Cheiridium museorum, Cheyletus eruditus, Cheyletus malaccensis, Tyrophagus putrescentiae, Withius subruber

Chao, Yung-Chang. 1954. Insects in Grain Elevators at Pullman and Albion, Washington.” Pan-Pacific Entomol. 30: 260–262. Scenopinus fenestralis (Omphrale fenestralis)

Chen, Chuan, Wei Yao, Yan Wang, and Jtanbin Wang. 1999. “The Capture Effect of Probe Traps on Stored-Product Insect Natural Enemies. p. 1492–1495. In Z. Jin, Q. .Liang, Y. Liang, X. Tan, and L. Guan eds. Proceedings of the 7th International Working Conference on Stored-Product Protection, Sichuan Publishing House of Science and Technology, Chengdu, China. Anisopteromalus calandrae, Lariophagus distinguendus

Chittenden, F. H. 1897. Parasites of Flour and Meal Moths (Illustrated). In Some little-known insects affecting stored vegetable products: a collection of articles detailing certain original observations make upon insects of the class. U. S. Bur. Entomol. Bulletin 8 – New Series: 38–43. Anisopteromalus calandrae, Apanteles carpatus, Apanteles ephestiae, Bracon brevicornis, Chremylus elaphus (Chremylus rubiginosus), Exochus mansuetor, Phytomyptera tarsalis (Clausicella tarsalis, Lispidea tarsalis), Habrobracon hebetor, Hemiteles tineae, Limnerium ephestiae, Melanophora roralis, Theocolax elegans (Chaetospila elegans), Theocolax formiciformis, Venturia canescens (Omorga frumentaria)

Cobanoglu, S. 1996. Determination, Distribution and Hosts of Acarina Group Harmful and Useful on Stored Products in the Province of Edirne (Turkey). Turkish Entomol. J. 20: 199–210. Acaropsellina sollers, Blattissocius tarsalis, Cheyletus eruditus, Cheyletus trouessarti, Tyrophagus putrescentiae

Coombs, C. W., and G. E. Woodroffe 1968. Changes in the Arthropod Fauna of an Experimental Bulk of Stored Wheat. Journal of Applied Ecology 5: 563–574. Cheyletus eruditus, Cheyletus trouessarti , Dibrachys cavus,  Lariophagus distinguendus, Lyctocoris campestris, Meteorus cespitator, Tyrophagus putrescentiae

Cornes, M. A. 1964. A Revised Listing of the Insects Associated with Stored Products in Nigeria. Ann. Rpt. Nigerian Stored Prod. Res. Instit. Techn. Bull.19: 96–119. Anisopteromalus calandrae, Antrocephalus mitys (Antrocephalus mahensis), Cephalonomia tarsalis, Cerocephala dinoderi, Dinarmus basalis (Dinarmus laticeps), Eupelmus vuilleti (Bruchocida vuilleti), Habrobracon hebetor, Holepyris glabratus (Holepyris hawaiiensis), Holepyris sylvanidis (Rhabdepyris zeae), Exkorynetes analis (Korynetes analis), Lariophagus dstinguendus, Lyctocoris campestris, Necrobia ruficollis, Necrobia rufipes, Peregrinator biannulipes, Plastanoxus munroi, Plastonoxus westwoodi, Reduvius minutus, Thaneroclerus buqueti, Xylocoris flavipes

Corpuz-Raros L, A., G. C. Sabio, and M. Velasco-Soriano. 1988. “Mites Associated with Stored Products, Poultry Houses and House Dust in the Philippines. Philippine Entomologist 7: 311–21. Acarapsis docta (Acaropsellina docta), Blattissocius dendriticus, Blattissocius keegani, Blattissocius tarsalis, Cheletomorpha lepidopterorum, Cheyletus fortis, Cheyletus malaccensis, Cunaxa capreolus, Tyrophagus putrescentiae

Costa,V.A., E.C. Guzzo, A.L. Lourencao, M.A.G.C. Tavares and J.D. Vendramim 2007. Occurrence of Dinarmus basalis in Callosobruchus analisin stored Soybean in Sao Paulo, Brazil. Scientia Agricola 64(3): 301-302. Dinarmus basalis

Cotterell, G. S. 1952. Insects associated with export produce in southern Nigeria. Bulletin of Entomological Research 43(1): 145–152. Antrocephalus mitys (Antrocephalus mahensis), Cephalonomia tarsalis, Habrobracon hebetor, Holepyris glabratus (Holepyris hawaiiensis), Holepyris sylvanidis (Rhabdepyris zeae), Lyctocoris campestris, Plastanoxus munroi

Cotton, Richard Thomas, and Newell Emanuel Good 1937. Annotated list of the insects and mites associated with stored grain and cereal products, and of their arthropod parasites and predators. USDA Miscellaneous Publication 258. Acarophenax tribolii, Anisopteromalus calandrae (Aplastomorpha calandrae), Anisopteromalus mollis, Antrocephalus mitys (Antrocephalus aethiopicus, Antrocephalus mahensis), Apanteles araeceri, Apanteles carpatus, Apanteles ephestiae (Apanteles nephoptericis), Bassus hawaiicola, Blattisocius tineivorus (Typhlodromus tineivorus), Bracon brevicornis, Carcinops pumilio (Carcinops quattuordecimstriata), Cephalonomia gallicola (Cephalonomia quadridentate, Cephalonomia xambeui). Cephalonomia hypobori (Cephalonomia nigricornis), Cephalonomia tarsalis (Cephalonomia carinata, Cephalonomia meridionalis), Cephalonomia watersoni, Cerocephala cornigera, Cerocephala dinoderi, Chelifer cancroides, Cheyletus eruditus (Cheyletus seminivorus), Chremylus elaphus (Chremylus rubiginosus), Clausicella floridensis (Epigrimyia floridensis), Cleonymus texanus (Ptinobius texanus), Cryptus ptinivorus, Dendrophilus punctatus, Dibrachys cavus, Dibrachys clisiocampae, Dimachus discolor,  Entedon longiventris, Eupelmus cushmani, Eupelmus javae, Eurytoma tylodermatis, Goniozus columbianus, Goniozus emigratus (Perisierola emigrate), Habritys brevicornis, Habrobracon hebetor, Hecabolus sulcatus, Hemiteles museorum, Hemiteles tineae, Holepyris glabratus (Holepyris hawaiiensis), Holepyris sylvanidis (Parepyris sylvanidis, Rhabdepyris zeae), Hypsicera curvator (Metacoelus mansuetor), Korynetes caeruleus (Corynetes coeruleus), Laelius anthrenivorus, Laelius centratus (Laelius trogodermatis), Laemostenus terricola (Aechmites terricola), Lariophagus distinguendus, Leucodesmia nigriventris, Lispe uliginosa (Helina uliginosa), Lyctocoris campestris, Melanophora roralis, Meraporus requisitus, Mesostenus gracilis, Nemeritis caudatula, Orius madeirensis, Pyemotes ventricosus (Pediculoides ventricosus), Philolema javensis (Aximopsis javensis). Philolema tephrosiae (Aximopsis tephrosiae), Phytomyptera tarsalis (Lispidea tarsalis), Plastanoxus chittendeni, Plastonoxus westwoodi, Platyspathius bisignathus (Spathius bisignatus), Pteromalus cerealellae (Habrocytus cerealellae), Pteromalus pyrophilus, Pteromalus tritici, Reduvius personatus, Saprinus semistriatus, Scenopinus fenestralis, Scenopinus glabrifrons, Sclerodermus domesticus, Sclerodermus immigrans, Spathius exarator, Thaneroclerus buqueti (Thaneroclerus girodi), Theocolax elegans (Chaetospila elegans), Theocolax formiciformis (Cephalonomia formiciformis), Tillus notatus, Trichogramma evanescens, Trichogramma minutum,Venturia canescens (Idechthis canescens), Xylocoris cursitans, Zatropis incertus

Davis, R. A. 1947. Notes on stored product pests in Burma and Singapore. Entomologist 29(1005): 36–40. Anisopteromalus calandrae (Aplastomorpha calandrae), Cephalonomia waterstoni, Dinarmus laticeps (Bruchobius laticeps), Habrobracon hebetor, Theocolax elegans (Chaetospila elegans)

Dhaliwal, GS, and GS Battu. 1976. Occurrence of Anisopteromalus calandrae (Howard) in Laboratory Culture of Rhizopertha dominica F. Bulletin of Grain Technology 14(2): 156–157.

Diamond, V. R. 1929. Biology of Nemeritis canescens, A parasite of the Mediterranean flour moth. Entomological Society of Ontario.Annual Report 60: 84–89. Venturia canescens (Nemeritis canescens)

Doane, R. W. 1919. Weevils in Australian wheat in California. Journal of Economic Entomology 12: 309–312. Anisopteromalus calandrae (Mesaporus calandrae)

Dolinski, M. G., W. Hanec, and S. R. Loschiavo 1971. Triticale as a New Host for Stored Grain Insects. The Manitoba Entomologist 5: 54. Cephalonomia waterstoni

Durrant, John Hartley. 1921. Insects Associated with Grain. Roy. Soc. of the Grain Pests (War) Committee 9(4): 33–52. Anisopteromalus calandrae (Aplastomorpha vandinei), Bracon brevicornis, Dinarmus basalis (Bruchobius laticeps), Holepyris sylvanidis (Rhabdepyris zeae), Lariophagus dstinguendus, Lariophagus puncticollis, Necrobia rufipes, Theocolax elegans (Choetospila elegans), Venturia canescens

El Bolok, M. M., I. I. Ismail, and H. A. Elshabrawy. 1990. Survey and Relative Abundance of Insects Attacking Onion in Field and Store with the Accompanied Natural Enemies at Giza and Assiut Regions. Annals of Agricultural Science, Moshtohor 28(3): 1799–1804. Cheyletus malaccensis, Tyrophagus putrescentiae, Xylocoris galactinus

Eliopoulos, P. A., C. G. Athanaaiou, and C. H. Buchelos 2002. Occurrence of Hymeopterous Parasitoids of Stored Product Pests in Greece. IOBC/WPRS Bulletin 25(3): 127–139. Anisopteromalus calandrae, Cephalonomia tarsalis, Cephalonomia waterstoni,Habrobracon (Bracon) hebetor,Heterospilus prosopidis, Holepyris hawaiensis,Holepyris sylvanidis, Lariophagus distinguendus, Pteromalus cerealellae, Theocolax elegans, Venturia canescens

Eliopoulos, P. A., Athanassiou, C. G., Palyvos, N. E., Stathas, G. J., and Buchelos, C. T. 2003. Dominance and frequency of predatory mites in stored products in Greece. Pages 258-262 in: Advances in Stored Product Protection. Proc. 8th.Int. Work. Conf. on Stored Prod. Prot., York, UK. Acarophenax tribolii, Acaropsis docta, Acaropsellina sollers (Acaropsis sollers), Blattisocius keegani, Blattisocius tarsalis, Cheletomorpha lepidopterum, Cheyletus aversor, Cheyletus eruditus, Cheyletus malaccensis, Cheyletus trouessarti, Cunaxa setirostris, Pyemotes ventricosus (Pediculoides ventricosus)

Eliopoulos, P. A., and C. G. Athanassiou 2004. Seasonal Occurrence of Dried Fig Pests and their Parasitoids in a Fig Warehouse in Greece. IOBC/WPRS Bulletin 27(9): 159–171. Cephalonomia tarsalis, Habrobracon hebetor, Holepyris silvanidis, Venturia canescens

El-Sayed, Ferial, and Mona Ghallab. 2007. Survey on mites associated with major insect pests infesting stored grains in Middle Delta. ACARINES: Journal of the Egyptian Society of Acarology 1(1): 29-38. Acaropsellina sollers, Apostigmaeus aegyptiacus, Blattisocius tarsalis, Blattisocius keegani, Cheyletus malaccensis, Cheyletus malayensis, Cheyletus fortis, Cunaxa capreolus, Nodele simplex, Pyemotes ventricosus, Spinibdella bifurcate, Spinibdella depressa  

El-Shafie, H.A.F. 2012. List of Arthropod Pests and Their Natural Enemies Identified Worldwide on Date Palm, Phoenix Dactylifera L. Agriculture and Biology Journal of North America 3(12): 516–524. Habrobracon hebetor, Parasierola swirskiana, Phanerotoma flavitestacea (Phanerotoma ocuralis), Venturia canescens, Xylocoris galactinus

Emam, A. K., H. E. Hanafy, S. I. Salama, and I. M. Badoor. 2013. Survey and Molecular Identification of Insect Pests Infesting Stored Leguminous Seeds and Their Associated Parasitoids. Journal of Applied Sciences Research 9(1): 928–36. Dinarmus basalis, Eupelmus vuilleti, Uscana lariophaga

Emmanouel, N. G., C. Th. Buchelos, and C.Th. E. Dukidis. 1994. A Survey on the Mites of Stored Grain in Greece. Journal of Stored Products Research 30(2): 175–178. Acaropsis docta, Androlaelaps casalis, Blattisocius keegani, Cheyletus eruditus, Pyemotes tritici, Tyrophagus putrescentiae

Fenilli, Renato 1978. Identificacao da entomofauna em armazens de soja em grao e de farelo no estado do Parana Brasil (Identification of entomofauna in warehouses of soybean and bran in the state of Parana Brazil). O Biologico 7: 161-164. Apanteles carpatus, Habrobracon hebetor, Peregrinator biannulipes

Froggatt, W. W. 1912. Parasitic enemies of the Mediterranean flour moth (Ephestia kuehniella, Zeller.). Agricultural Gazette of New South Wales 23: 307–311. Habrobracon hebetor, Venturia canescens

Gates, Michael William. 1992. Population Dynamics of Lesser Grain Borer, Rusty Grain Beetle, and Cephalonomia waterstoni in Commercial Elevators. Oklahoma State University.

Giles, P. H. 1964. Insect infestation of sorghum stored in granaries in northern Nigeria. Bulletin of Entomological Research 55 (3): 573–588. Exkorynetes analis (Korynetes analis), Reduvius minutus, Xylocoris flavipes

Gothilf, S. 1969. Natural enemies of the carob moth Ectomyelois ceratoniae (Zeller). Entomophaga, 14: 195–202. Anisopteromalus mollis, Antrocephalus mitys, Apanteles lacteus, Bracon brevicornis, Clausicella suturata, Perilampus tristis, Phanerotoma flavitestacea, Pristomerus vulnerator

Guertin, D. S., P. J. Ode, M. R. Strand, and M. F. Antolin. 1996. Host-Searching and Mating in an Outbreeding Parasitoid Wasp. Ecological Entomology 21(1): 27–33. Habrobracon hebetor

Habibpour, B, Kamali, K. and Meidani, J. 2002. Insects and mites associated with stored products and their arthropod parasites and predators in Khuzestan province (Iran). Integrated Protec Stored Prod, IOBC Bull.25(3): 89–91. Acarapsis docta, Cheyletus malaccensis, Habrobracon breviconis, Habrobracon hebetor, Hypoaspis aculeifer, Hypoaspis sclerotarsa, Proctolaelaps pygmaeus, Theocolax elegans (Choetospila elegans)

Hagstrum D.W. 1987. Seasonal variation of stored wheat environment and insect populations. Environmental Entomology 16: 77–83. Cephalonomia waterstoni

Hagstrum, D. W., and Sharp, J. E. 1975. Population studies on Cadra cautella in a citrus pulp warehouse with particular reference to diapause. J. Econ. Entomol. 68: 11-14. Blattisocius tarsalis, Habrobracon hebetor

Haines, C. P. 1999. Arthropod natural enemies in stored products-overlooked and under-exploited. p. 1205-1226. Proc. 7th Int. Work. Conf. Stored-product Protection, Beijing China,Sichuan Publishing House of Science and Technology, China. Acarophenax tribolii, Androlaelaps casalis, Anisopterornalus calandrae, Blattisocius dentriticus, Blattisocius keegani, Blattisocius tarsalis, Carcinops troglodytes, Cephalonomia tarsalis, Cephalonomia waterstoni, Cheyletus fortis, Cheyletus malaccensis, Cunaxa setirostrus, Euchalcidia caryobori, Dinarmus basalis,Habrobracon hebetor, Holepyris hawaiiensis, Holepyris sylvanidis, Hypocacculus metallescens, Lariophagus distinguendus,Laemotmetus rhizophagoides, Lyphia orientalis, Peregrinator biannulipes, Plastanoxus munroi, Plastanoxus westwoodi,Pronematus bachewinqi, Thaneroclerus buqueti, Theocolax elegans,Tydeus australensis, Withius piger, Xylocoris flavipes

Hameed, A. A., A. A. Al-Taweel, H. K. Al-Zubaidy, and A. M. Tark. 2011. Fields and Warehouses Survey for Seasonal Occurrence of Date Moth Ephestia Spp. and Their Parasitoid Bracon hebetor in Some Orchards and Warehouses in Iraq. Iraqi J. Agric. Res. 16(2): 96–108.

Hameed, A. A., A. A. Al-Taweel, A. J. Eass, R. N. Shalal, and Z. A. Al-Garbawi. 2010. Surveying Ephestia Spp. and the Parasitoid Bracon hebetor in Date Palm Orchards and Date Stores of Five Governorates in Iraq.Acta Horticulturae, ISHS 882: 985–93.

Hamlin, John Calhoun, William Doyle Reed, and M. E. Phillips 1931. Biology of the Indian-meal moth on dried fruits in California. USDA Technical Bulletin 242. Clausicella floridensis (Drepanoglossa floridensis), Habrobracon hebetor, Venturia canescens

Harish, G., A. Naganagoud, A. G. Sreenivas, Sharangouda Hiregoudar, and B. Kisan 2018. Diversity and Distribution of Stored Grain Insect Pests of Pulses and their Natural Enemies. Indian Journal of Agricultural Research 52(2): 157–161. Dinarmus basalis, Dinarmus vagabundus

Hayashi, T., S. Nakamura, P. Visarathanonth, J. Uraichuen, and R. Kengkanpanich. 2004. Stored Rice Insect Pests and Their Natural Enemies in Thailand. Book, Whole. Funny Publishing Co. Ltd., Bangkok, Thailand: JIRCAS Interantional Agricultural Series. No. 13 Anisopteromalus calandrae, Antrocephalus mitys, Cephalonomis tarsalis, Cerocephala dinoderi, Habrobracon hebetor, Holepyris sylvanidis, Lariophagus distinguendus, Proconura caryobori, Proconura minusa, Theocolax elegans

Hayhurst, H. 1937. Insect infestation of stored products. Annals of Applied Biology 24(4): 797–807. Habrobracon hebetor

Helbig, J. 1998. Ability of Naturally Occurring Parasitoids to Suppress the Introduced Pest Prostephanus truncatus (Horn) (Coleoptera, Bostrichidae) in Traditional Maize Stores in Togo. Journal of Stored Products Research 34(4): 287–295. Anisopteromalus calandrae, Cephalonomia tarsalis, Cephalonomia waterstoni, Holepyris sylvanidis, Pediobius furvus, Plastonoxus westwoodi, Theocolax elegans

Hirabayashi, Jan Misa. 2004. Dynamics and Parasitism of Stored Product Pests in Traditionally Stored Maize in Rural Mexico. PhD diss., University of California, Berkeley. Anisopteromalus calandrae, Lariophagus distinguendus, Pteromalus cerealellae

Horton, P.M., 1982. Stored product insects collected from on-farm storage in South Carolina. Journal of the Georgia Entomological Society (USA) 17: 485-491. Cardiastethus assimilis, Cephalonomia tarsalis, Cheletomorpha lepidopterum, Cheyletus eruditus

Hossaein MA, Ali AT, Hashem K, Somayeh K. 2009. Stored product pest and their parasitoid wasps in Mashhad, Iran. Adv Env Biol.3(3): 239–243. Habrobracon hebetor, Cephalonomia tarsalis, Anisopteromalus calandrae, Theocolax elegans, Venturia canescens

Hussain, S., and N. A. Aslam. 1970. Some Observations on a Beneficial Reduviid Bug, Amphibolus venator Klug (Hemiptera: Reduviidae). J. Agric. Pakistan 21(1): 37–42.

Imamura, Taro, Akihiro Miyanoshita, Satoshi Furui, Kazushi Miyatake, Yuka Hirai, Yuko Tai and Hiroshi Nakakita 2014. Investigation of stored-product insect pests and their natural enemies by traps baited with brown rice in grain drying and conditioning facilities and grain warehouses. Rep. Nat’l Food Res. Inst 78: 1-9.  Anisopteromalus calandrae, Lariophagus distinguendus

Islam, W., K. M. Ahmed, and M. A. M. Malek. 1985. Hymenopterous Parasites on Insect Pests of Two Stored Pulses. Bangladesh Journal of Zoology 13(2): 81–85. Anisopteromalus calandrae, Dinarmus basalis, Habrobracon hebetor, Theocolax elegans (Choetospila elegans)

Johnson, J.A., K.A. Valero, M.M. Hannel, and R.F. Gill 2000. Seasonal Occurrence of Postharvest Dried Fruit Insects and their Parasitoids in a Culled Fig Warehouse. Journal of Economic Entomology 93(4): 1380–1390. Anisopteromalus calandrae,Cerchysiella utilis, Cephalonomia gallicola, Cephalonomiatarsalis,Cephalomia waterstoni, Habrobracon hebetor, Holepyris sylvanidis, Laelius centratus, Mesostenus gracilis, Plastonoxus westwoodi, Psilochalcis brevialata,Venturia canescens

Kapil, R. P., and J. P. Chaudhary. 1975. Record of New Hymenopterous Parasites from Khapra Beetle, Trogoderma granarium Everts (Coleoptera: Dermestidae). Indian Journal of Entomology 35(4): 353–55. Anisopteromalus calandrae, Dinarmus laticeps

Katiyar, R. R. et al. 1981. New record of Scenopinus indicus (Scenopinidae: Diptera) as a predator on stored grain pests in India. Bulletin of Grain Technology 19(2): 140.

Kiruba, Solomon, Sarma Mohan, Sathiadas Sam Manohar Das, and Smaragdi Papadopoulou 2012. Experimental Confirmation of the Bruchidae Natural Parasitism Efficacy Using an Innovative Device, Friendly to the Environment (Parasitoid Facilitator Bin). Biotechnology and Biotechnological Equipment 26(1): 2722–2725. Dinarmus basalis, Uscana lariophaga

Kishani-Farahani, H., S. H. Goldansaz, and Q. Sabahi. 2012. A Survey on the Overwintering Larval Parasitoids of Ectomyelois Ceratoniae in Three Regions in Iran. Crop Protection 36: 52–57. Apanteles myeloenta, Campoplex tumidulus, Habrobracon hebetor, Temelucha decorata, Venturia canescens

Ketoh, GK, AI Glitho, and J. Huignard. 2002. Susceptibility of the Bruchid Callosobruchus Maculatus (Coleoptera : Bruchidae) and Its Parasitoid Dinarmus basalis (Hymenoptera : Pteromalidae) to Three Essential Oils. Journal of Economic Entomology 95(1): 174–182.

Key, K. H. L., and I. F. B. Common. 1959. Observations on the ecology of the clothes moths Tineola bisselliella (Humm.) and Tinea pellionella L. in a bulk wool store. Australian Journal of Zoology 7(1): 39–77. Apanteles carpatus, Chremylus elaphus (Chremylus rubiginosus)

Khan, Butool Ali and Anwarullah, M. 1970. Predators and parasites associated with stored grain pests in Karachi, West Pakistan. Science and Industry 7(1-2): 45–49. Anisopteromalus calandrae, Habrobracon hebetor, Theocolax elegans

Kiritani, K. 1957. Investigation on the faunal compositions in stored products: II. The faunal composition in relation to kinds of stored products. Applied Entomology and Zoology Nihon Oyo Dobutsu Konchu Gakkai 1: 8–14. Habrobracon hebetor

Kiritani, Keizi 1958. On the distribution and seasonal prevalence of stored grain insects in a farm premises. Botyu-Kagaku 23: 164-172. Anisopteromalus calandrae (Neocatolaccus mamezophagus), Habrobracon hebetor, Lariophagus distinguendus

Konishi, Kazuhiko, Yoshinobu Nakatani, Kazuo Ogata, Porntip Visarathanonth and Satoshi Nakamura. 2006. Fauna of Natural Enemies of Stored Rice Pests in Thailand. Japan International Research Center for Agricultural Sciences (JIRCAS), Tsukuba, Japan Working Report 45: 31-37. Amphiareus constrictus, Amphibolus venator, Anisopteromalus calandrae, Antrocephalus mitys, Cardiastethus pygmaeus, Cephalonomia tarsalis, Cerocephala dinoderi, Habrobracon hebetor, Holepyris sylvanidis, Joppeicus paradoxus, Lariophagus dstinguendus, Peregrinator biannulipes, Proconura caryobori, Proconura minusa, Thaneroclerus buqueti, Theocolax elegans, Xylocoris flavipes

Ksentini, Ines, Juan Carlos Monje, Taieb Jardak, and Najiba Zeghal 2010. Naturally occurring egg parasitoids of the genus Trichogramma (Hymenoptera: Trichogrammatidae) in a pomegranate orchard in Tunisia. Entomological science 13(1): 99-106. Ectomyelois ceratoniae by Trichogramma bourarachae, Trichogramma cacoeciae, Trichogramma evanescens, Trichogramma oleae

Kundra, S. 1976. Some Observations on Biology of Dinarmus vagabundus Timberlake (Pteromalidae-Hymenoptera). Bulletin of Grain Technology 14(2): 126–129. Found in laboratory culture Callosobruchus maculatus

Li, Zhaohui, Zheng Fangqiang, Ye Baohua, Liu Guilin, Li Qiang and Li Haiping 1999. Bionomics of Lariophagus distinguendus (Foerster) (Hymenoptera: Pteromalidae) and its control effectiveness to Maize weevils. p. 1258-1260. In  Jin, Z.; Liang, Q.; Liang, Y.; Tan, X.; and Guan, L. (Eds.), Proceedings of the 7th International Working Conference on Stored-Product Protection, 14-19 October 1998, Beijing, China. Sichuan Publishing House of Science and Technology, Chengdu, China

Lim, J., M. Oh, J. Lee, and S. Lee. 2007. Cephalonomia gallicola (Hymenoptera: Bethylidae), New to Korea, an ectoparasitoid of the cigarette beetle, Lasioderma serricorne. J. Asia-Pac Entomol. 10: 335-338.

Lin, T. 1968. A Survey on the Granary Insects and Their Natural Enemies in Taiwan  (in Chinese). J. Agric. Res. China 17: 39–45. Anisopteromalus calandrae (Neocatolaccus mamezophagus), Habrobracon hebetor, Lariophagus distinguendus, Pyemotes ventricosus (Pediculoides ventricosus), Pteromalus cerealellae, (Habrocytus cerealellae), Theocolax elegans(Chaetospila elegans), Venturia canescens

Livingstone, E. M., and W. D. Reed. 1936. Insect Fauna of Cured Tobacco in Storage in the United States. Journal of Economic Entomology 29(5): 1017–1022. Mesostenus gracilis, Venturia canescens

Lotfalizadeh, H., Hossenini, F. 2013. A survey of storage pests parasitoids (Hymenoptera) in Iran. Journal of Egypt University Faculty of Agriculture, 1: 113-120. Anisopteromalus calandrae, Cephalonomia tarsalis, Bracon vrevicornis, Dinarmus vagabundus, Habrobracon hebetor, Laelius anthrenivorus, Lariophagus distinguendus, Theocolax elegans, Theocolax formiciformis

Lukas, J., and V. Stejskal. 2010. Liotryphon Punctulatus (Ratzeburg, 1848) (Hymenoptera: Ichneumonidae) – a Parasitoid of Ephestia kuehniella Larvae. p. 726–728. In Proceedings of the 10th International Working Conference on Stored Product Protection edited by M. O. Carvalho, P. G. Fields, C. S. Adler, F. H. Arthur, C. G. Athanassiou, J. F. Campbell, F. Fleurat-Lessard, et al., Julius Kühn-Institut. Habrobracon hebetor, Venturia canescens

Lukáš, J., E. Žd’árková, M. Wakefield, and J. Hubert 2002. Parasitoids Occurring in Food-Processing Factories and Grain Stores. p. 83–86. In Eva Zdarkova, Mauren Wakefield, Jan Lukas, and Jan Hubert eds. Biological Control of pest insects and mites with special reference of entomophthorales. Proceedings of the second meeting of working group 4, COST Action 842,  Research Institute of Crop Production, Prague 30-31 May 2002 Anisopteromalus calandrae, Cephalonomia tarsalis, Habrobracon hebetor, Venturia canescens.

Markham, R. H. 1981.The Ecology of Insect Populations in Maize Storage Cribs in Nigeria. PhD diss., University of London. Antrocephalus crassipes, Antrocephalus mitys, Anisopteromalus calandrae, Bruohocida vuilleti, Cephalononria formiciformis, Cerocephala dinoderi, Eupelmus urozonus, Habobracon hebetor, Habrocytus cerealellae, Holepyris hawaiiensis, Plastanoxus westwoodi Rhabdepyris zea, Theocolax elegans, Zeteticontus laevigatus (parasite Cathartus quadricollis)

Maurya, K. R., B. Dev, and Z. Jamil. 1984. Population Patterns of Androlaelaps casalis (Acarina Laelapinae) in Stored Grains from Lucknow (India). Biological Memoirs 9 (2): 151–154.

McNally, Alfred Gordon 1932. The biology of Lariophagus distinguendus Foerster a Hymenopterous parasite of the grain and rice weevils. MA thesis, University of Western Ontario, Canada

McFarlane, J. A. 1963. Annotated record of Coleoptera, Lepidoptera, Hemiptera and Hymenoptera associated with stored produce in Jamaica. Tropical Agriculture 40(3): 211–216. Amphibolus venator, Anisopteromalus calandrae, Blattisocius dentriticus (Melichares dentriticus), Plastanoxus munroi (Cheyletus munroi), Habrobracon hebetor, Holepyris sylvanidis (Rhabdepyris zeae), Lyctocoris campestris, Necrobia rufipes, Thaneroclerus buqueti, Xylocoris flavipes

Mital, V. P. 1969. Anisopteromalus calandrae, Howord. (Hymenoptera: Chalcicidae) A new record of pupal parasite of groundnut bruchid (Caryedon gonagra, F.) A serious pest of groundnut (Arachis hypogea L.) and Tamarind (Tamarindus indica L.). Bulletin of Grain Technology 7(4): 234–235.

Mohamed, Azza A., Seham A. Ezz El-Dein, and Tawfik A. Tawfik. 2019. Survey of main mites and insect species associated with stored maize in Giza governorate. Journal of Plant Protection and Pathology 10(5): 257-260. Androlaelaps casalis, Blattisocius tarsalis, Cheyletus eruditus, Cheyletus malaccensis, Tyrophagus putrescentiae

Mohan-Rao, H. N., A. N. Ramesh, S. Kulkarni, and M. S. Rao. 1973. A survey on storage infestation in Bombay warehouses. Bulletin of Grain Technology 11(3/4): 222-224 Anisopteromalus calandrae, Cerocephala dinoderi, Dinarmas vagabundus, Habrobracon hebetor, Lariophagus distinguendus, Theocolax elegans

Monge, Jean Paul, and Jacques Huignard. 1991. Population fluctuations of two bruchid species Callosobruchus maculatus (F.) and Bruchidius atrolineatus (Pic) (Coleoptera Bruchidae) and their parasitoids Dinarmus basalis (Rondani) and Eupelmus vuilleti (Crawford) (Hymenoptera, Pteromalidae, Eupelmidae) in a storage situation in Niger. Journal of African Zoology Revue de Zoologie Africaine 105: 187–196.

Munro, James Watson and W. S. Thomson 1929. Report on insect infestation of stored cocao. Empire Marketing Board (E. M. B.) 24, London. Habrobracon hebetor (Microbracon hebetor, Holepyris hawaiiensis

Myers, John Golding 1929. Notes on some natural enemies of Plodia interpunctella and Silvanus surinamensis in Australia. Bulletin of Entomological Research 20: 425–430. Habrobracon hebetor, Venturia canescens

Nahar, S. C., and S. K. Gupta. 1980. Preliminary Report of Stored Grain Mites of Bihar. Bulletin of Grain Technology 18(2): 130–134. Acarophenax tribolii, Cheyletus erudius, Cheyletus fortis, Tyrophagus putrescentiae

O’Connor, James P., and C. Ronayne. 2002. Irish Records of Holepyris sylvanidis (Brèthes) and Plastanoxus munroi Richards (Hym., Bethylidae). Entomologists Monthly Magazine 138: 62

O’Farrell, A. F., and P. M. Butler. 1936. Insects and mites associated with the storage and manufacture of foodstuffs in Northern Ireland. Economic Proceedings, Royal Dublin Society 3(22): 343–407. Acaropsellina docta (Acaropsis docta), Anisopteromalus calandrae, Cephalonomia tarsalis, Cephalonomia waterstoni, Cheletomorpha lepidopterorum (Cheyletus venustissimus), Cheyletus carnifex, Cheyletus eruditus, Cheyletus trouessarti, Dimachus discolor, Eucheyletia flabellifera (Cheyletus flabellifera),  Habrobracon hebetor, Lyctocoris campestris, Necrobia rufipes, Necrobia violacea, Plastanoxus munroi (Cheyletus munroi), Pteromalus cerealella (Habrocytus cerealella), Spathius exarator, Theocolax elegans (Choetopsila elegans), Theocolax formiciformis, Venturia canescens (Nemeritis canescens), Xylocoris flavipes

Padmanaban, B. 1997. Incidence of pteromalid parasitoids on stored rice in Arunachal Pradesh.
Indian Journal of Hill Farming 10(1/2): 94-95. Cerocephala dinoderi, Theocolax elegans

Parker, B. L., R. H. Booth, and C. P. Haines. 1981. Arthropods infesting stored cassava (Manihot esculenta Crantz) in peninsular Malaysia. Protection Ecology 3: 141–156. Anisopteromalus calandrae, Theocolax elegans

Parker, W. B. 1915. Control of dried-fruit insects in California. USDA Bull. 235. Habrobracon hebetor

Pascual-Villalobos, M.J., R. Carreres, J. Riudavets, M. Aguilar, J.M. Bozal, M. C. García, and P. del Estal 2006. Plagas Del Arroz Almacenado y Sus Enemigos Naturales En Espana (Pests of Stored Rice and their Natural Enemies in Spain). Bol. San. Veg. Plagas 32: 225–231.

Pascual Villalobos, M. J., Baz, A. and Del Estal, P. 2005. Occurrence of psocids and natural predators on organic rice in Calasparra (Murcia, Spain). J. Stored Prod. Res. 41: 231-235. Cheletomorpha lepidopterorum, Cheyletus malaccensis, Withius piger

Pereira, P. 1998. The Use of Pheromone Traps for Monitoring Ephestia kuehniella Zeller (Lepidoptera: Pyralidae) and Detection of Parasitoids in Flour Mills. IOBC Bulletin 21 (3): 111–117.Anisopteromalus calandrae, Bracon brevicornis, Venturia canescens

Pérez-Benavides, A. Lucía, Fernando Hernández-Baz, Jorge M. González, Jesús Romero-Nápoles, Paul E. Hanson, and Alejandro Zaldívar-Riverón. 2020. Integrative taxonomy to assess the species richness of chalcidoid parasitoids (Hymenoptera) associated to Bruchinae (Coleoptera: Chrysomelidae) from Mexico. Revista mexicana de biodiversidad 91, e913492 Brasema neococcidis, Chryseida bennetti, Eurydinoteloides incerta, Horismenus butcheri, Horismenus depressus, Horismenus missouriensis, Pteromalus fuscipennis, Torymus moazopi  – Acanthoscelides obtectus and/or Acanthoscelides obvelatus on Phaseolus vulgaris and/or Phaseolus lunatus

Prevett P. F. 1961 Field infestation of cowpea (Vigna unguiculata)pods by beetles of the families Bruchidae and Curculionidae in Northern Nigeria. Bull. Entomol. Res.52, 635-645. Anisopteromalus calandrae, Bruchocida vuilleti

Proctor, D. L. 1970. Annotated list of arthropods associated with stored food products in Zambia. Tropical Products Institute L18. Anisoptermalus calandrae, Antrocephalus mitys (Antrocephalus mahensis), Cheyletus malaccensis, Habrobracon hebetor, Holepyris sylvanidis (Rhabdepyris zeae), Lariophagus distinguendus, Peregrinator biannulipes, Theocolax elegans (Chaetospila elegans), Tyrophagus putrescentiae, Xylocoris flavipes

Pulpan, J., and P.H. Verner. 1965. Control of Tyroglyphoid Mites in Stored Grain by Predatory Mite Cheyletus eruditus (Schrank). Canadian Journal of Zoology 43(3): 417–432.

Rawnsley, J. 1959. The natural relationship between the hyrnenopterous parasite, Bracon hebetor, and its host, the cocoa moth Ephestia cautella; The importance of this relationship in designing chemical control. Ghana Cocoa Mark. Bd., Insect Control Unit, Pub. 7, p. 1-3.

Rawnsley, J. 1968. Biological studies in Ghana in the control of Cadra cautella (Wlk.), the tropical warehouse moth. Ghana J. Agr. Sci., 1(2):155-159. Habrobracon hebetor

Reed, C.R., D.W. Hagstrum, P.W. Flinn, and R.F. Allen 2003. Wheat in Bins and Discharge Spouts, and Grain Residues on Floors of Empty Bins in Concrete Grain Elevators as Habitats for Stored-Grain Beetles and their Natural Enemies. Journal of Economic Entomology 96(3): 996–1004. Anisopteromalus calandrae, Cephalonomia waterstonii, Habrobracon hebetor, Theocolax elegans

Richards, O. W. and G. V. B. Herford 1930. Insects found associated with cacao, spices and dried fruits in London warehouses. Ann. Appl. Biol. 17: 367-395. Cephalonomia tarsalis, Habrobracon hebetor, Holepyris glabratus (Holepyris hawaiiensis), Lyctocoris campestris, Xylocoris flavipes (Piezostethus flavipes)

Richards, O. W., and W. S. Thomson. 1932. A Contribution to the Study of the Genera Ephestia, Gn.(including Strymax, Dyar), and Plodia, Gn.(Lepidoptera, Phyeitidae), with Notes on Parasites of the Larvae. Transactions of the Royal Entomological Society of London 80(2): 169-250 Habrobracon hebetor, Mesostenus gracilis, Venturia canescens

Richards, Owain Westmacott, and Nadia Waloff. 1946. Study of a Population of Ephestia elutella Hubner (Lep., Phycitidae) Living on Bulk Grain. Transactions of the Royal Entomological Society of London 97(11): 253–298. Diadegma chrysostictos (Angitia chrysosticta, Horogenes chrysostictos), Habrobracon hebetor, Venturia canescens

Richards, Owain Westmacott  1949. Parasitic Hymenoptera found in British houses, warehouses, and ships. I. Ichneumonidae. Proceedings of the Royal Entomological Society of London 18(1-2): 19–35. Diadegma chrysostictos (Angitia chrysostictos), Gelis cinctus (Hemiteles cinctus), Hypsicera curvator (Metacoelus mansuetor), Xenolytus bitinctus (Phygadeuon bitinctus), Venturia canescens (Nemeritis canescens)

Riudavets, J., É. Lucas, and M. José Pons. 2002. Insects and Mites of Stored Products in the Northeast of Spain. IOBC/Wprs Bulletin 25(3): 41–44. Anisopteromalus calandrae, Blattisocius tarsalis, Lariophagus distinguendus, Tyrophagus putrescentiae

Runner, George Alfred 1919. The tobacco beetle: an important pest in tobacco products. USDA Bulletin 737. Thaneroelerus girodi, Anisopteromalus calandrae (Aplastomorpha pratti, Aplastomorpha vandinei)

Ryoo, M. I., H. W. Cho, and Y. B. Kim. 1990. Ecological Successions of Arthropod Communities in Stored Rough Rice, Polished Rice and Brown Rice. Korean. Korean J.Appl.Entomol. 29: 31–42. Anisopteromalus calandrae, Lariophagus distinguendus, Pymotes tritici

Saleh, Alaa. 2012. Ecological and Biological Study of the Cigarette Beetle, Lasioderma Serricorne and Survey Its Natural Enemies. MS thesis, Faculty of Agriculture, Damascus University, Syria. Amblyseius swirskii, Anisopteromalus calandrae, Ericydnus sipylus, Lariophagus distinguendus, Pteromalus cereallae, Tyrophagus putrescentiae

Salmond, K. F. 1957. Investigations into Grain Storage Problems in Nyasaland Protectorate with Special Reference to Maize (Zea Mays L.). Colon. Res. Publ. (London, H.M.S.O.) 21: 1–49. Cheyletomorpha lepidoptorum (Cheyletomorpha venustissima), Plastanoxus munroi (Cheyletus munroi), Stenowithius bayoni (Stenowithius ugandanus), Xylocoris afer (Piezostethus afer), Xylocoris flavipes

Savidan-Neiderer, A. 2002. Tritrophic Interactions in Maize Storage Systems. Ph.D. Thesis, Université de Neuchâtel, Neuchâtel, Switzerland. Anisopteromalus calandrae, Lariophagus dstinguendus, Pteromalus cerealellae

Schneider, Heinrich, Christian Borgemeister, Mamoudou Sétamou, Hippolyte Affognon, Albert Bell, Matthias E. Zweigert, Hans-Michael Poehling, and Fritz Schulthess. 2004. Biological Control of the Larger Grain Borer Prostephanus truncatus (Horn) (Coleoptera : Bostrichidae) by Its Predator Teretrius nigrescens (Lewis) (Coleoptera : Histeridae) in Togo and Benin. Biological Control 30(2): 241–55.

Schneider, V., Beukeboom, L.W., Driessen, G., Lapchin, L., Bernstein, C. and Van Alphen, J.J.M. 2002. Geographical distribution and genetic relatedness of sympatrical thelytokous and arrhenotokous populations of the parasitoid Venturia canescens (Hymenoptera). Journal of Evolutionary Biology 15: 191–200.

Sedlacek, J.D.; Price, B.D.; Sharkey, M.J.; Hill, S.J. and Weston, P.A. 1998. Parasitoids found in on-farm shelled corn in Kentucky. J. Agric. Entomol. 15: 223–230. Anisopteromalus calandrae, Cephalonomia tarsalis, Cephalonomia waterstoni, Habrobracon hebetor, Theocolax elegans

Seini, Seidu Shaibu. 1998. A search for indigenous Parasitoids for the control of the larger grain borer, Prostephanus truncatus (Horn)(Coleoptera: Bostrichidae). MS thesis, University of Ghana. Anisopteromalus calandrae, Dinarmus basalis, Eupelmus vuilleti, Holepyris sylvanidis found cowpea and maize purchased from market.

Sharma, Sanjay Kumar and M. Ashfaque Ali 1991. Activity of natural enemies against Callosobruchus maculatus (F.) in store houses of pigeonpea. Indian Journal of Zoological Spectrum 2(1-2): 31-34. Dinarmus basalis

Shen, Chao-Peng 1984. List and survey of stored-product mites in China. p. 680–686. In Proceedings of the 3rd  International Working Conference on Stored-Product Entomology edited by R. B. Mills, V. F. Wright, J. R. Pedersen, W. H. McGaughey, R. W. Beeman, K. J. Kramer, R. D. Speirs, and C. L. Storey, Kansas State University. Acarophenax tribolii, Acaropsellina sollers (Acaropsis sollers), Androlaelaps casalis, Blattisocius dentriticus, Blattissocius tarsalis, Blattisocius keegani, Cheletomorpha lepidopterorum, Cheyletus eruditus, Cheyletus malaccensis, Cheyletus trouessarti, Cunaxa setirostris, Haemogamasus pontiger, Tyrophagus putrescentiae

Showell, Harry 1928. A Useful Parasite of the Dried Fruit Moth. Journal of Agriculture of South Australia 31(11): 1048–1056. Habrobracon hebetor (Habrobracon juglandis)

Simmons, Perez, William Doyle Reed, and E. A. McGregor. 1931. Fig insects in California. USDA Circ. 157. Cephalonomia tarsalis, Clausicella floridensis (Drepanoglossa floridensis), Habrobracon hebetor (Habrobracon juglandis), Venturia canescens (Nemeritis canescens)

Souza, Moises S. de, Cesar A. D. Teixeira, Celso O. Azevedo, Valmir A. Costa, and Jose N. M. Costa 2006. Occurrence of Cephalonomia stephanoderis Betrem (Hymenoptera : Bethylidae) in Coffee Plantations of the Brazilian Amazonia. Neotropical Entomology 35(4): 560–562.

Stein-Beling, I. 1969. Uber den ausflug der schlupfwespe Nemeritis canescens Grav. und uber die bedeutung des geruchssinnes bei der ruckkehr zum wirt (The Flight of the Parasite Nemeritis canescens and the Significance of the Olfactory Organs for the Return). Biologisches Zentralblatt 54: 147–169.

Stejskal, V., J. Lukáš, E. Žd’árková, M. Wakefield, and J. Hubert 2002. Spatial Arrangement of Venturia canescens and Ephestia kuehniella in the Extremely Infested Pasta-Producing Factory: A Case History. 116–117. In Biological Control of pest insects and mites with special reference of entomophthorales. Proceedings of the second meeting of working group 4, COST Action 842 , Research Institute of Crop Production.

Stejskal, V., P. Kosina, and L. Kanyomeka. 2006. Arthropod Pests and their Natural Enemies in Stored Crops in Northern Namibia. Journal of Pest Science 79(1): 51–55. Blattissocius tarsalis, Cephalonomia waterstoni, Habrobracon hebetor

Suma, P., Amante, M., Bella, S., La Pergola, A. and Russo, A. 2014. Stored-product insect natural enemies in wheat industry in Sicily. IOBC-WPRS Bulletin 98: 227–233.

Tadesse, A. 1996. Insects and Other Arthropods Recorded from Stored Maize in Western Ethiopia. African Crop Science Journal 4(3): 339–343. Anisopteromalus calandrae, Holepyris sylvanidis, Theocolax elegans

Takahashi, K., T. Ohbayashi, and N. Sota 2000. Investigation of Stored-Product Insect Pests and their Natural Enemies in Chichijima Island, Ogasawara (Bonin), Japan [In Japanese with English Summary]. Jpn.J.Ent.(N.S.) 3: 97–103. Carcinop pumilio, Fulvius anthocoroides

Takahashi, K., and M. V. Romero 2001. Investigation of Stored-Product Insect Pests and their Natural Enemies in Okinawa-Jima and Ishigaki-Jima Islands, Okinawa, Japan. Japanese Journal of Entomology (New Series) 4(3): 91–97. Venturia canescens

Tawfik, M. F., K. T. Awadallah, and M. M. H. Abdella 1984. Population Studies on Wheat Grain Insect Pests and their Natural Enemies at Four Localities in Egypt. Arab Gulf Journal of Scientific Research 2(1): 267–280. Anisopteromalus calandrae, Holepyris hawaiiensis, Lariophagus distinguendus, Peregrinator biannulipes (Allaeocranum biannulipes), Theocolax elegans, Xylocoris flavipes

Tawfik, M. F. S., K. T. Awadallah, M. M. El-Husseini, and A. I. Afifi 1984. Survey on Stored Drug Insect Mite Pests and their Associated Natural Enemies in Egypt. Bulletin of the Entomological Society of Egypt 65: 267–274. Acaropsis docta, Anisopteromalus calandrae, Blattisocius tarsalis, Cheyletus malaccensis, Cunaxa capreolus,, Lariophagus distinguendus, Peregrinator biannulipes (Allaeocranum biannulipes), Termatophylum insigne, Xylocoris flavipes

Tigar, BJ, PE Osborne, GE Key, ME Flores, and M. Vazquez. 1994. Distribution and Abundance of Prostephanus truncatus (Coleoptera, Bostrichidae) and Its Predator Teretriosoma nigrescens (Coleoptera, Histeridae) in Mexico. Bulletin of Entomological Research 84(4): 555–565.

van Alebeek, F. A. N. 1996. Natural suppression of bruchid pests in stored cowpea (Vigna unguiculata (L.) Walp.) in West Africa. Int. J. Pest Manage. 42: 55-60. Dinarmus spp., Eupelimus spp., Xylocoris flavipes

Walker, D. J. 1979. Insects Associated with Stored Maize and Other Harvested Food Crops in Swaziland. Journal of the Entomological Society of Southern Africa 42(2): 331–335. Anisopteromalus calandrae, Habrobracon hebetor, Holepyris sylvanidis (Rhabdepyris zeae), Peregrinator biannulipes, Theocolax elegans (Chaetospila elegans), Xylocoris afer

Walker, D. J., and R. A. Boxall. 1974. Annotated List of the Insects Associated with Stored Products in Ethiopia, Including Notes on Mites Found in Harar Province. East African Agricultural and Forestry Journal 39(3): 330–335. Anisopteromalus calandrae, Cheyletus eruditus, Habrobracon hebetor, Lariophagus distinguendus, Necrobia rufipes, Necrobia violacaea, Theocolax elegans (Chaetospila elegans), Xylocoris afer

Waller, J. B. 1982. A Record of Venturia canescens (Hymenoptera, Ichneumonidae) in New Zealand. New Zealand Entomologist 7(3): 319.

Waloff, N., and O. W. Richards. 1946. Observations on the Behaviour of Ephestia elutella Hubner (Lepidoptera, Pycitidae Breeding on Bulk Grain. Transactions of the Royal Entomological Society of London 97(12): 299–335. Angitia chrysostica, Habrobracon hebetor, Venturia canescens

Waterston, J. 1921. Report on Parasitic Hymenoptera, bred from pests of stored grain. Reports of the Grains Pests (War) Committee, Royal Society 9: 8–32. Dinarmus basalis (Bruchobius laticeps), Holepyris sylvanidis (Rhabdepyris zeae), Lariophagus distinguendus, Theocolax elegans (Chaetospila elegans)

Werner, Petr, Zuzana Kucerova, Vaclav Stejskal, and Eva Žďárková. 1999. “Stored Product Pests in Stored Grain in the Czech Republic. p. 64–68. In Proceedings of the 7th International Working Conference on Stored-Product Protection, edited by Z. Jin, Q. Liang, Y. Liang, X. Tan, and L. Guan, Sichuan Publishing House of Science and Technology. Cheyletus eruditus, Tyrophagus putrescentiae

Williams, R. N., Floyd, E. H. 1971: Effect of two parasites, Anisopteromalus calandrae and Choetospila elegans upon populations of the maize weevil under laboratory and natural conditions. J. Econ. Entomol. 64: 1407-1408.

Xu, Zaifu, Liqiong Weng, and Junhua He. 2008. A New Species of Cephalonomia Westwood (Hymenoptera : Bethylidae) Parasitizing Rhizopertha dominica (F.) (Coleoptera : Bostrychidae). Proceedings of the Entomological Society of Washington 110(2): 477–480. Cephalonomia rhizoperthae

Yoshida, Toshiharu and Kenzi Kawano 1959. Fauna and community structure of the insects in the grain stored at farm-houses: The ecological studies of the pests infesting stored grains. part 3. Miyazaki Daigaku Gakugei Gakubu Kiyo III Shizen Kagaku Memoirs of the Faculty of Liberal Arts and Education, Miyazaki University; Natural Science 7: 33–61. Anisopteromalus calandrae (Neocatolaccus mamezophagus), Habrobracon hebetor, Lariophagus distinguendus, Theocolax elegans, Trichomalus apanteloctenus

Yoshida, T. and Kenzi Kawano 1958. Seasonal fluctuation of the number of insects in the grains stored at farmhouse, part 1. The ecological studies of the pests infesting stored grains. part 2. Miyazaki Daigaku Gakugei Gakubu Kiyo III Shizen Kagaku Memoirs of the Faculty of Liberal Arts and Education, Miyazaki University; Natural Science 5: 11–23. Anisopteromalus calandrae (Neocatolaccus mamezophagus), Lariophagus distinguendus, Theocolax elegans, Trichomalus apanteloctenus

Zdarkova, Eva. 1967. Stored Food Mites in Czechoslovakia. Journal of Stored Product Research 3: 155–175. Acaropsis docta, Acaropsellina sollers (Acaropsis sollers), Androlaelaps casalis (Haemolaelaps casalis),  Blattisocius keegani, Blattisocius tarsalis, Cheyletomorpha lepidoptorum, Cheyletus eruditus, Cheyletus malaccensis, Cheyletus trouessarti,  Cheyletus aversor, Haemogamasus pontiger, Tyrophagus putrescentiae

Zipagan, M. B., and E. P. Pacumbaba 1996. Insect Pests of Copra and Copra Meal in the Philippines and their Natural Enemies. Technical Reports Philippine Coconut Authority